The beloved Galápagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has died. Zoo officials say she died on Nov. 20. Gramma arrived at the zoo ...

Gramma the tortoise, believed to be 141 years old and the oldest resident of the San Diego Zoo, was euthanized after ...

Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of age.

The San Diego Zoo announced Gramma, the Galápagos tortoise, died on Nov. 20. She was about 141 years old based on estimates ...

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